Jones has called up 33 players to assemble today at Pennyhill Park ahead of England’s first of two matches against Wales in build-up to the Rugby World Cup in Japan.

With no new players added to the squad following England’s 12-day training camp in Treviso, Italy, Mike Brown (Harlequins), Alex Dombrandt (Harlequins), Ben Spencer (Saracens) and Ben Te’o (unattached) are not included in this week’s squad.

Jones said: “We have worked hard and now move into a four-game preparation phase where the development of the team tactically is paramount. We are grateful for the strong opposition to test our game fitness.

“By the time we fly out on the 8th of September we will be ready to win the Rugby World Cup. We are not there yet but we have four games to get ready.

“Players excluded from being involved in this camp are naturally disappointed but may get another opportunity so they must be ready.“

England will play Wales (11 August) and Ireland (24 August) at Twickenham Stadium and Italy (6 September) in Newcastle as part of the Quilter Internationals as well as an away fixture against Wales (17 August) in Cardiff.

England will begin their Rugby World Cup campaign in Japan against Tonga in Sapporo (22 September, KO 11:15am UK time) before playing USA Rugby in Kobe (26 September, KO 11:45am UK time), Argentina in Tokyo (5 October, KO 09:00am UK time) and France in Yokohama (12 October, KO 09:15am UK time).

England squad

Forwards
Dan Cole (Leicester Tigers, 85 caps)
Luke Cowan-Dickie (Exeter Chiefs, 11 caps)
Tom Curry (Sale Sharks, 10 caps)
Charlie Ewels (Bath Rugby, 10 caps)
Ellis Genge (Leicester Tigers, 9 caps)
Jamie George (Saracens, 37 caps)
Maro Itoje (Saracens, 27 caps)
George Kruis (Saracens, 32 caps)
Joe Launchbury (Wasps, 58 caps)
Courtney Lawes (Northampton Saints, 71 caps)
Lewis Ludlam (Northampton Saints, uncapped)
Joe Marler (Harlequins, 59 caps)
Kyle Sinckler (Harlequins, 22 caps)
Jack Singleton (Saracens, uncapped)
Sam Underhill (Bath Rugby, 9 caps)
Billy Vunipola (Saracens, 41 caps)
Mako Vunipola (Saracens, 53 caps)
Harry Williams (Exeter Chiefs, 17 caps)
Mark Wilson (Newcastle Falcons / Sale Sharks, 13 caps)

Backs
Joe Cokanasiga (Bath Rugby, 4 caps)
Elliot Daly (Saracens, 30 caps)
Owen Farrell (Saracens, 70 caps)
George Ford (Leicester Tigers, 55 caps)
Piers Francis (Northampton Saints, 4 caps)
Willi Heinz (Gloucester Rugby, uncapped)
Jonathan Joseph (Bath Rugby, 40 caps)
Joe Marchant (Harlequins, uncapped)
Jonny May (Leicester Tigers, 45 caps)
Ruaridh McConnochie (Bath Rugby, uncapped)
Henry Slade (Exeter Chiefs, 22 caps)
Manu Tuilagi (Leicester Tigers, 32 caps)
Anthony Watson (Bath Rugby, 33 caps)
Ben Youngs (Leicester Tigers, 85 caps)

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